Many good suggestions, keep them coming. Let me add what I discovered at the boat today. On the starboard engine the level came back up a bit, so I think the changes in reservoir levels on that engine are just normal expansion and contraction, perhaps also filling some trapped air.
The port engine however, is a different story. I found a lot of coolant under the engine, sitting in the engine bilge, enough to account for the loss from the reservoir. This is the engine that serves the hot water heater, but I could see no evidence of leaks from those hoses or fittings. Also, the coolant in the engine bilge must have come from the engine, since the expansion reservoir is located outside this engine bilge, so the expansion tank did not leak it out. I removed the pressure cap from the metal reservoir on top of the engine, and could not feel any coolant with my fingers down to 4” or so.
With this new info, any ideas? Neither engine got hot during my trip, although we had some heavy seas one day. Is the spring loaded cap on the metal reservoir suspect?